South American Indians who, at the time of the Spanish conquest in 1532, ruled an empire that extended along the Pacific coast and Andean highlands from the northern border of modern Ecuador to the Maule River in central Chile. The Inca established their capital at Cuzco (Peru) in the 12th century. They began their conquests in the early 15th century and within 100 years had gained control of an Andean population of about 12,000,000 people. These Quechua-speaking tribes' origins are uncertain. Their vast empire had a centralized organization and at its head was the ruler, 'Son of the Sun', worshipped as a god in his own lifetime. As a divine king he was above the law, and as a despotic ruler he was very much the political head of the state. Administration was in the hands of officials drawn from the Inca nobility and from the chiefs of conquered tribes. An efficient road system, along which relays of messengers could travel 250 km in a day, ensured that Cuzco was kept informed of developments all over the empire. These same roads allowed Inca forces to be quickly moved into any province which showed signs of rebellion. This centralization was both the strength and the weakness of the Inca state. The unifying force was the ruler in person, and the death of Huayna Capac precipitated a crisis. Civil war broke out when two of his sons, Huascar and Atahuallpa, disputed the succession. Atahuallpa won the war, but before he could consolidate his position he was seized and murdered by Francisco Pizarro's Spaniards in 1532. Without a leader the Inca system could not function. Most of the empire was quickly brought under Spanish control, but an independent Inca group held out in the Urubamba valley until 1572. Viracocha Inca was the creator, culture hero, and supreme deity of the Inca, but the religion embraced a pantheon of gods of nature. The most actively worshipped were the sun and, by extension, the emperor, who was considered the son of the sun. The Temple of the Sun, built at the pre-Incan ceremonial center of Pachacamac suggests some incorporation of earlier religions. Archaeologically, the Inca culture is characterized by fine quality stone masonry, agricultural terraces, mass-produced and standardized pottery forms (aryballus), and metal objects. The considerable architectural skill of the Inca is reflected in Cyclopean masonry, although many buildings were constructed using rectangular dressed stone blocks as well as adobe. The basic dwelling-unit was a cluster of single rooms arranged around a rectangular courtyard and was most often enclosed by a wall. Writing was unknown, but the quipu was used for keeping records. Agriculture was based on plant foods, especially potato, manioc, quinoa, and maize. Domesticated animals included dog, llama, cava (guinea pig), and alpaca. Fine textiles were woven using a simple backstrap loom. The civilization was the largest and most powerful political unit in all the prehistoric America. It has been argued that the whole of Inca achievement relied heavily on a variety of political, societal and religious infrastructures already in place before their ascendancy.